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Fauna Sinica Insecta Vol.75 Coleoptera Histeroidea Sphaeritidae, Synteliidae and Histeridae，The present work studies the taxonomy of the superfamily Histeroidea of China. Histeroidea is a very important superfamily of the Polyphaga beetles in the order Coleoptera and represents a special clade of Staphyliniformia. Histeroidea includes now three families, namely, Sphaeritidae, Synteliidae and Histeridae; last one is very speciose of 4400 species worldwide. Most histerid beetles are predatory, saproxylic or xylophagy, living on some fungi or decaying materials of plants and (or) animals; part are also as myrmecophilus, living together with ants. Many studies focused on evolution and phylogeny of Histeridae; some entomologists want to find out how to use histerid beetles and natural enemies in pest control; ecologists tried to use these beetle species as bio-indicators in dead wood decaying and forest management.

This book, as one of the series published under the name "Fauna Sinica", is a comprehensive study of taxonomy and systematic of Chinese Histeroidea and includes the results of our long-termed studies on this beetle group. Based on the taxonomic revision, we included in this work all the known species of the superfamily Histeroidea recorded in China up to 2017. The exact fauna data are 3 families (Sphaeritidae, Synteliidae and Histeridae)67 genera 303 species. The Histeridae is a very large taxon comparing to the other two families and is composed of 10 specieose subfamilies. This family takes the major part of Chinese histeroid fauna. This work includes keys to species or other higher taxa, descriptions of all families, subfamilies, genera and almost all species found in China. All species and higher taxa are revised with a complete citations; messages of specimens observed are also included; species geographical distributions are also up-dated. Morphological characters are displayed in 228 figures and a total of ca. 2000 ling drawing; these may benefit the identification for the readers. In a very brief review text, this work includes also the following contents: a perspective to the taxonomic studies, the morphology of adults, eggs, larvae and pupae; systematic and its changes in the taxonomy of histerid history; geographical distributions in the different regions of China; other messages of the biological and ecological features, etc. Of course this work includes also the new species and revisions published by my team members.

 

目录

前言

总论 1

一、国内外研究概况 1

二、形态特征 3

（一）成虫 3

（二）幼虫 23

（三）蛹 26

（四）卵 27

三、分类发展历史和系统发育关系 27

（一）阎甲总科的系统地位 27

（二）阎甲总科的分类 28

（三）扁圆甲科的系统地位 29

（四）长阎甲科的系统地位 29

（五）阎甲科的分类发展历史和系统发育关系 30

四、地理分布 37

（一）属的分布类型及其特点 37

（二）种的分布类型及其特点 39

五、生物学和生态学特性 52

（一）食性和栖息环境 52

（二）世代和发育历期 55

（三）成虫的生活习性和行为学 56

（四）天敌 58

六、经济意义 58

各论 60

阎甲总科Histeroidea Gyllenhal, 1808 60

一、扁圆甲科Sphaeritidae Shuckard, 1839 60

1. 扁圆甲属Sphaerites Duftschmid, 1805 61

(1) 双色扁圆甲Sphaerites dimidiatus Jurecek, 1934 61

(2) 黑扁圆甲Sphaerites glabratus (Fabricius, 1792) 63

(3) 翔扁圆甲Sphaerites involatilis Gusakov, 2004 64

(4) 亮扁圆甲Sphaerites nitidus L？bl, 1996 64

(5) 暗扁圆甲Sphaerites opacus L？bl et Háva, 2002 66

(6) 孔扁圆甲Sphaerites perforatus Gusakov, 2017 67

二、长阎甲科Synteliidae Lewis, 1882 67

2. 长阎甲属Syntelia Westwood, 1864 68

(7) 大卫长阎甲Syntelia davidis Fairmaire, 1889 68

(8) 马氏长阎甲Syntelia mazuri Zhou, 2003 69

(9) 中华长阎甲Syntelia sinica Zhou, 2003 71

三、阎甲科Histeridae Gyllenhal, 1808 72

（一）细阎甲亚科Niponiinae Fowler, 1912 74

3. 细阎甲属Niponius Lewis, 1885 74

(10) 沟细阎甲Niponius canalicollis Lewis, 1901 75

(11) 凹细阎甲Niponius impressicollis Lewis, 1885 76

(12) 姬细阎甲Niponius osorioceps Lewis, 1885 77

(13) 山崎细阎甲Niponius yamasakii Miwa, 1934 79

（二）嗜蚁阎甲亚科Chlamydopsinae Bickhardt, 1914 81

4. 奇异阎甲属Eucurtiopsis Silvestri, 1926 81

(14) 奇异阎甲Eucurtiopsis mirabilis Silvestri, 1926 82

（三）脊阎甲亚科Onthophilinae MacLeay, 1819 82

5. 脊阎甲属Onthophilus Leach, 1817 83

(15) 黄角脊阎甲Onthophilus flavicornis Lewis, 1884 85

(16) 细脊阎甲Onthophilus foveipennis Lewis, 1885 86

(17) 丽江脊阎甲Onthophilus lijiangensis Zhou et Luo, 2001 88

(18) 原脊阎甲Onthophilus ordinarius Lewis, 1879 90

(19) 粗脊阎甲Onthophilus ostreatus Lewis, 1879 91

(20) 席氏脊阎甲Onthophilus silvae Lewis, 1884 93

(21) 斯氏脊阎甲Onthophilus smetanai Mazur, 1994 94

(22) 瘤脊阎甲Onthophilus tuberculatus Lewis, 1892 95

6. 毛脊阎甲属Epiechinus Lewis, 1891 95

(23) 多刺毛脊阎甲Epiechinus hispidus (Paykull, 1811) 96

(24) 马氏毛脊阎甲Epiechinus marseuli Lewis, 1900 97

(25) 南洋毛脊阎甲Epiechinus taprobanae Lewis, 1892 98

(26) 树毛脊阎甲Epiechinus arboreus (Lewis, 1884) 99

（四）小阎甲亚科Tribalinae Bickhardt, 1914 100

7. 悦阎甲属Epierus Erichson, 1834 100

(27) 索氏悦阎甲Epierus sauteri Bickhardt, 1913 101

8. 副悦阎甲属Parepierus Bickhardt, 1913 101

(28) 中国副悦阎甲Parepierus chinensis Zhang et Zhou, 2007 102

(29) 异刺副悦阎甲Parepierus inaequispinus Zhang et Zhou, 2007 104

(30) 刘氏副悦阎甲Parepierus lewisi Bickhardt, 1913 106

(31) 梳刺副悦阎甲Parepierus pectinispinus Zhang et Zhou, 2007 107

9. 小阎甲属Tribalus Erichson, 1834 109

1) 小阎甲亚属Tribalus Erichson, 1834 110

(32) 点小阎甲Tribalus (Tribalus) punctillatus Bickhardt, 1913 110

2) 真小阎甲亚属Eutribalus Bickhardt, 1921 111

(33) 鸽小阎甲Tribalus (Eutribalus) colombius Marseul, 1864 111

(34) 柯氏小阎甲Tribalus (Eutribalus) koenigius Marseul, 1864 113

(35) 欧氏小阎甲Tribalus (Eutribalus) ogieri Marseul, 1864 115

（五）阎甲亚科Histerinae Gyllenhal, 1808 115

一）突胸阎甲族Exosternini Bickhardt, 1914 116

10. 短臀阎甲属Anaglymma Lewis, 1894 117

(36) 环短臀阎甲Anaglymma circularis (Marseul, 1864) 117

11. 圆臀阎甲属Notodoma Lacordaire, 1854 119

(37) 蕈圆臀阎甲Notodoma fungorum Lewis, 1884 120

12. 凹背阎甲属Epitoxus Lewis, 1900 122

(38) 亚洲凹背阎甲Epitoxus asiaticus Vienna, 1986 123

(39) 布拉凹背阎甲Epitoxus bullatus (Marseul, 1870) 123

13. 长臀阎甲属Cypturus Erichson, 1834 124

(40) 埃长臀阎甲Cypturus aenescens Erichson, 1834 125

14. 缘尾阎甲属Paratropus Gerstaecker, 1867 127

(41) 坎达拉缘尾阎甲Paratropus khandalensis Kanaar, 1997 127

二）扁阎甲族Hololeptini Hope, 1840 129

15. 扁阎甲属Hololepta Paykull, 1811 130

3) 扁阎甲亚属Hololepta Paykull, 1811 131

(42) 乌苏里扁阎甲Hololepta (Hololepta) amurensis Reitter, 1879 131

(43) 鲍氏扁阎甲Hololepta (Hololepta) baulnyi Marseul, 1857 133

(44) 扁阎甲Hololepta (Hololepta) depressa Lewis, 1884 135

(45) 长扁阎甲Hololepta (Hololepta) elongata Erichson, 1834 137

(46) 费氏扁阎甲Hololepta (Hololepta) feae Lewis, 1892 138

(47) 希氏扁阎甲Hololepta (Hololepta) higoniae Lewis, 1894 140

(48) 印度扁阎甲Hololepta (Hololepta) indica Erichson, 1834 141

(49) 亮扁阎甲Hololepta (Hololepta) laevigata Guérin-Ménéville, 1833 143

(50) 尼泊尔扁阎甲Hololepta (Hololepta) nepalensis Lewis, 1910 145

(51) 钝扁阎甲Hololepta (Hololepta) obtusipes Marseul, 1864 145

(52) 平扁阎甲Hololepta (Hololepta) plana (Sulzer, 1776) 147

三）方阎甲族Platysomatini Bickhardt, 1914 148

16. 巨颚阎甲属Megagnathos Penati et Zhang, 2009 150

(53) 巨颚阎甲Megagnathos lagardei Vienna et Ratto, 2015 151

17. 沟颚阎甲属Silinus Lewis, 1907 151

(54) 沟颚阎甲Silinus procerus (Lewis, 1911) 151

18. 长卵阎甲属Platylister Lewis, 1892 153

4) 长卵阎甲亚属Platylister Lewis, 1892 154

(55) 黑长卵阎甲Platylister (Platylister) atratus (Erichson, 1834) 154

(56) 缅甸长卵阎甲Platylister (Platylister) birmanus (Marseul, 1861) 156

(57) 柬埔寨长卵阎甲Platylister (Platylister) cambodjensis (Marseul, 1864) 158

(58) 霍氏长卵阎甲Platylister (Platylister) horni (Bickhardt, 1913) 161

(59) 平氏长卵阎甲Platylister (Platylister) pini (Lewis, 1884) 162

(60) 缝长卵阎甲Platylister (Platylister) suturalis (Lewis, 1888) 164

(61) 凯氏长卵阎甲Platylister (Platylister) cathayi Lewis, 1900 165

5) 平尾阎甲亚属Popinus Mazur, 1999 166

(62) 孔氏长卵阎甲Platylister (Popinus) confucii (Marseul, 1857) 166

(63) 筛臀长卵阎甲Platylister (Popinus) dahdah (Marseul, 1861) 168

(64) 光亮长卵阎甲Platylister (Popinus) lucillus (Lewis, 1891) 169

(65) 独长卵阎甲Platylister (Popinus) unicus (Bickhardt, 1912) 171

19. 阔咽阎甲属Apobletes Marseul, 1860 173

(66) 缘阔咽阎甲Apobletes marginicollis Lewis, 1888 173

(67) 绍氏阔咽阎甲Apobletes schaumei Marseul, 1860 175

20. 方阎甲属Platysoma Leach, 1817 178

6) 方阎甲亚属Platysoma Leach, 1817 179

(68) 贝氏方阎甲Platysoma (Platysoma) beybienkoi Kryzhanovskij, 1972 179

(69) 短线方阎甲Platysoma (Platysoma) brevistriatum Lewis, 1888 179

(70) 中国方阎甲Platysoma (Platysoma) chinense Lewis, 1894 181

(71) 平方阎甲Platysoma (Platysoma) deplanatum (Gyllenhal, 1808) 183

(72) 达氏方阎甲Platysoma (Platysoma) dufali Marseul, 1864 185

(73) 重方阎甲Platysoma (Platysoma) gemellun (Cooman, 1929) 187

(74) 朝鲜方阎甲Platysoma (Platysoma) koreanum Mazur, 1999 189

(75) 滑方阎甲Platysoma (Platysoma) rasile Lewis, 1884 189

(76) 裂方阎甲Platysoma (Platysoma) rimarium Erichson, 1834 189

(77) 四川方阎甲Platysoma (Platysoma) sichuanum Mazur, 2007 189

(78) 井上方阎甲Platysoma (Platysoma) takehikoi ？hara, 1986 191

7) 长方阎甲亚属Cylister Cooman, 1941 192

(79) 狭方阎甲Platysoma (Cylister) angustatum (Hoffmann, 1803) 193

(80) 长方阎甲Platysoma (Cylister) elongatum (Thunberg, 1787) 195

(81) 线方阎甲Platysoma (Cylister) lineare Erichson, 1834 197

(82) 细方阎甲Platysoma (Cylister) lineicolle Marseul, 1873 199

(83) 云南方阎甲Platysoma (Cylister) yunnanum (Kryzhanovskij, 1972) 201

21. 近方阎甲属Niposoma Mazur, 1999 201

(84) 刘氏近方阎甲Niposoma lewisi (Marseul, 1873) 202

(85) 申氏近方阎甲Niposoma schenklingi (Bickhardt, 1913) 204

(86) 斯氏近方阎甲Niposoma stackelbergi (Kryzhanovskij, 1976) 205

(87) 台湾近方阎甲Niposoma taiwanum (Hisamatsu, 1965) 206

22. 卡那阎甲属Kanaarister Mazur, 1999 207

(88) 阿萨姆卡那阎甲Kanaarister assamensis (Lewis, 1900) 207

(89) 隐卡那阎甲Kanaarister celatum (Lewis, 1884) 208

(90) 库氏卡那阎甲Kanaarister coomani (Thérond, 1955) comb. nov. 209

23. 似真卵阎甲属Eurosomides Newton, 2015 211

(91) 微似真卵阎甲Eurosomides minor (Rossi, 1792) 211

24. 沟尾阎甲属Liopygus Lewis, 1891 213

(92) 沟尾阎甲Liopygus andrewesi Lewis, 1906 214

25. 短卵阎甲属Eblisia Lewis, 1889 216

8) 短卵阎甲亚属Eblisia Lewis, 1889 217

(93) 田舍短卵阎甲Eblisia (Eblisia) pagana Lewis, 1902 217

(94) 小短卵阎甲Eblisia (Eblisia) pygmaea (Bickhardt, 1913) 218

(95) 索氏短卵阎甲Eblisia (Eblisia) sauteri (Bickhardt, 1912) 219

(96) 苏门答腊短卵阎甲Eblisia (Eblisia) sumatrana (Bickhardt, 1912) 221

9) 点尾阎甲亚属Chronus Lewis, 1914 222

(97) 卡西短卵阎甲Eblisia (Chronus) calceata (Cooman, 1931) 222

26. 真卵阎甲属Eurylister Bickhardt, 1920 224

(98) 萨氏真卵阎甲Eurylister satzumae (Lewis, 1899) 225

(99) 树真卵阎甲Eurylister silvestre (Schmidt, 1897) 226

27. 直沟阎甲属Mendelius Lewis, 1908 228

(100) 细直沟阎甲Mendelius tenuipes (Lewis, 1905) 228

28. 完额阎甲属Aulacosternus Marseul, 1853 229

(101) 完额阎甲Aulacosternus zelandicus Marseul, 1853 229

29. 大阎甲属Plaesius Erichson, 1834 231

10) 大阎甲亚属Plaesius Erichson, 1834 232

(102) 爪哇大阎甲Plaesius (Plaesius) javanus Erichson, 1834 232

(103) 莫氏大阎甲Plaesius (Plaesius) mohouti Lewis, 1879 234

11) 异胸阎甲亚属Hyposolenus Lewis, 1907 236

(104) 孟加拉大阎甲Plaesius (Hyposolenus) bengalensis Lewis, 1906 236

四）斜臀阎甲族Omalodini Kryzhanovskij, 1972 238

30. 脊额阎甲属Lewisister Bickhardt, 1912 239

(105) 脊额阎甲Lewisister excellens Bickhardt, 1912 239

五）阎甲族Histerini Gyllenhal, 1808 241

31. 歧阎甲属Margarinotus Marseul, 1853 242

12) 光折阎甲亚属Ptomister Houlbert et Monnot, 1923 244

(106) 阿葛歧阎甲Margarinotus (Ptomister) agnatus (Lewis, 1884) 245

(107) 阿若歧阎甲Margarinotus (Ptomister) arrosor (Bickhardt, 1920) 247

(108) 康夫歧阎甲Margarinotus (Ptomister) babai ？hara, 1999 247

(109) 博氏歧阎甲Margarinotus (Ptomister) boleti (Lewis, 1884) 249

(110) 尸生歧阎甲Margarinotus (Ptomister) cadavericola (Bickhardt, 1920) 251

(111) 海拉尔歧阎甲Margarinotus (Ptomister) hailar Wenzel, 1944 253

(112) 隐歧阎甲Margarinotus (Ptomister) incognitus (Marseul, 1854) 254

(113) 科氏歧阎甲Margarinotus (Ptomister) koltzei (Schmidt, 1889) 256

(114) 多齿歧阎甲Margarinotus (Ptomister) multidens (Schmidt, 1889) 257

(115) 大泽歧阎甲Margarinotus (Ptomister) osawai ？hara, 1999 259

(116) 理氏歧阎甲Margarinotus (Ptomister) reichardti Kryzhanovskij et Reichardt, 1976 261

(117) 纹歧阎甲指名亚种Margarinotus (Ptomister) striola striola (Sahlberg, 1819) 263

(118) 缝连歧阎甲Margarinotus (Ptomister) sutus (Lewis, 1884) 265

(119) 三线歧阎甲Margarinotus (Ptomister) tristriatus Wenzel, 1944 266

(120) 温氏歧阎甲Margarinotus (Ptomister) wenzelisnus Kryzhanovskij et Reichardt, 1976 266

(121) 魏氏歧阎甲Margarinotus (Ptomister) weymarni Wenzel, 1944 268

13) 毛折阎甲亚属Eucalohister Reitter, 1909 270

(122) 双斑歧阎甲Margarinotus (Eucalohister) bipustulatus (Schrank, 1781) 270

(123) 美斑歧阎甲Margarinotus (Eucalohister) gratiosus (Mannerheim, 1852) 272

14) 宽胫阎甲亚属Stenister Reichardt, 1926 274

(124) 暗歧阎甲Margarinotus (Stenister) obscurus (Kugelann, 1792) 274

15) 端线阎甲亚属Paralister Bickhardt, 1917 277

(125) 柯氏歧阎甲Margarinotus (Paralister) koenigi (Schmidt, 1888) 277

(126) 中亚歧阎甲Margarinotus (Paralister) laevifossa (Schmidt, 1890) 279

(127) 长圆歧阎甲Margarinotus (Paralister) oblongulus (Schmidt, 1892) 281

(128) 周歧阎甲Margarinotus (Paralister) periphaerus Mazur, 2003 282

(129) 暗红歧阎甲Margarinotus (Paralister) purpurascens (Herbst, 1792) 284

16) 普胫阎甲亚属Grammostethus Lewis, 1906 286

(130) 缅甸歧阎甲Margarinotus (Grammostethus) birmanus Lundgren, 1991 287

(131) 台湾歧阎甲Margarinotus (Grammostethus) formosanus Ohara, 1999 287

(132) 脆歧阎甲Margarinotus (Grammostethus) fragosus (Lewis, 1892) 289

(133) 勤歧阎甲Margarinotus (Grammostethus) impiger (Lewis, 1905) 291

(134) 日本歧阎甲Margarinotus (Grammostethus) niponicus (Lewis, 1895) 292

(135) 海西歧阎甲Margarinotus (Grammostethus) occidentalis (Lewis, 1885) 294

(136) 施氏歧阎甲Margarinotus (Grammostethus) schneideri Kapler, 1996 296

(137) 粪歧阎甲Margarinotus (Grammostethus) stercoriger (Marseul, 1880) 297

(138) 台岛歧阎甲Margarinotus (Grammostethus) taiwanus Mazur, 2008 299

17) 缝歧阎甲亚属Asterister Desbordes, 1920 299

(139) 完缝歧阎甲Margarinotus (Asterister) curvicollis (Bickhardt, 1913) 299

32. 突唇阎甲属Pachylister Lewis, 1904 301

18) 突唇阎甲亚属Pachylister Lewis, 1904 302

(140) 斯里兰卡突唇阎甲宽臀亚种Pachylister (Pachylister) ceylanus pygidialis Lewis, 1906 302

(141) 泥突唇阎甲Pachylister (Pachylister) lutarius (Erichson, 1834) 304

19) 沟唇阎甲亚属Sulcignathos Mazur, 2010 306

(142) 拙突唇阎甲Pachylister (Sulcignathos) scaevola (Erichson, 1834) 306

33. 完折阎甲属Nasaltus Mazur et W？grzynowicz, 2008 306

(143) 中国完折阎甲Nasaltus chinensis (Quensel, 1806) 307

(144) 东方完折阎甲Nasaltus orientalis (Paykull, 1811) 310

34. 阎甲属Hister Linnaeus, 1758 312

(145) 比斯阎甲Hister bissexstriatus Fabricius, 1801 314

(146) 同色阎甲Hister concolor Lewis, 1884 316

(147) 康吉阎甲Hister congener Schmidt, 1885 317

(148) 遥阎甲Hister distans Fischer de Waldheim, 1823 320

(149) 幻异阎甲Hister inexspectatus Desbordes, 1923 322

(150) 日本阎甲Hister japonicus Marseul, 1854 322

(151) 爪哇阎甲Hister javanicus Paykull, 1811 324

(152) 马氏阎甲Hister mazuri Kapler, 1997 326

(153) 巨爪阎甲Hister megalonyx Reichardt, 1922 328

(154) 普然阎甲Hister pransus Lewis, 1892 330

(155) 点足阎甲Hister punctifemur Mazur, 2010 332

(156) 糙阎甲东部亚种Hister salebrosus subsolanus (Newton, 1991) 332

(157) 谢氏阎甲Hister sedakovii Marseul, 1861 334

(158) 上海阎甲Hister shanghaicus Marseul, 1861 337

(159) 西伯利亚阎甲Hister sibiricus Marseul, 1854 339

(160) 简胸阎甲Hister simplicisternus Lewis, 1879 341

(161) 斯普阎甲Hister spurius Marseul, 1861 342

(162) 西藏阎甲Hister thibetanus Marseul, 1857 344

(163) 单色阎甲累氏亚种Hister unicolor leonhardi Bickhardt, 1910 346

35. 分阎甲属Merohister Reitter, 1909 348

(164) 吉氏分阎甲Merohister jekeli (Marseul, 1857) 349

36. 新植阎甲属Neosantalus Kryzhanovskij, 1972 351

(165) 新植阎甲Neosantalus latitibius (Marseul, 1861) 352

37. 三线阎甲属Eudiplister Reitter, 1909 355

(166) 米氏三线阎甲Eudiplister muelleri Kryzhanovskij et Reichardt, 1976 355

(167) 平坦三线阎甲Eudiplister planulus (Ménétries, 1848) 356

38. 清亮阎甲属Atholus Thomson, 1859 358

(168) 奇额清亮阎甲Atholus bifrons (Marseul, 1854) 359

(169) 双斑清亮阎甲Atholus bimaculatus (Linnaeus, 1758) 360

(170) 青色清亮阎甲Atholus coelestis (Marseul, 1857) 360

(171) 缘清亮阎甲Atholus confinis (Erichson, 1834) 362

(172) 窝胸清亮阎甲Atholus depistor (Marseul, 1873) 362

(173) 十二纹清亮阎甲十四纹亚种Atholus duodecimstriatus quatuordecimstriatus (Gyllenhal, 1808) 365

(174) 勒清亮阎甲Atholus levis Mazur, 2015 367

(175) 菲律宾清亮阎甲Atholus philippinensis (Marseul, 1854) 367

(176) 皮瑞清亮阎甲Atholus pirithous (Marseul, 1873) 369

(177) 纹尾清亮阎甲Atholus striatipennis (Lewis, 1892) 372

(178) 扭清亮阎甲Atholus torquatus (Marseul, 1854) 373

39. 毛腹阎甲属Asiaster Cooman, 1948 376

(179) 卡卡毛腹阎甲Asiaster calcator Cooman, 1948 376

(180) 库氏毛腹阎甲Asiaster cooteri Kapler, 1999 378

(181) 赫氏毛腹阎甲Asiaster hlavaci Lackner, 2004 379

（六）伴阎甲亚科Hetaeriinae Marseul, 1857 380

40. 伴阎甲属Hetaerius Erichson, 1834 381

(182) 悦伴阎甲Hetaerius optatus Lewis, 1884 381

（七）卵阎甲亚科Dendrophilinae Reitter, 1909 383

六）卵阎甲族Dendrophilini Reitter, 1909 384

41. 卵阎甲属Dendrophilus Leach, 1817 384

20) 卵阎甲亚属Dendrophilus Leach, 1817 385

(183) 宽卵阎甲Dendrophilus (Dendrophilus) xavieri Marseul, 1873 385

21) 拟卵阎甲亚属Dendrophilopsis Schmidt, 1890 387

(184) 展卵阎甲Dendrophilus (Dendrophilopsis) proditor (Reichardt, 1925) 387

七）小齿阎甲族Bacaniini Kryzhanovskij, 1976 389

42. 穆勒阎甲属Mullerister Cooman, 1936 389

(185) 日本穆勒阎甲Mullerister niponicus (Lewis, 1879) 389

(186) 东京湾穆勒阎甲Mullerister tonkinensis (Cooman, 1936) 391

43. 小齿阎甲属Bacanius LeConte, 1853 391

22) 小齿阎甲亚属Bacanius LeConte, 1853 392

(187) 科氏小齿阎甲Bacanius (Bacanius) collettei Gomy, 1999 392

(188) 卡氏小齿阎甲Bacanius (Bacanius) kapleri Gomy, 1999 394

(189) 天皇小齿阎甲Bacanius (Bacanius) mikado (Lewis, 1892) 395

44. 显臀阎甲属Abraeomorphus Reitter, 1886 397

(190) 台湾显臀阎甲Abraeomorphus formosanus (Hisamatsu, 1965) 397

八）丽尾阎甲族Paromalini Reitter, 1909 399

45. 簇点阎甲属Coomanister Kryzhanovskij, 1972 400

(191) 斯氏簇点阎甲Coomanister scolyti Mazur, 2007 400

46. 匀点阎甲属Carcinops Marseul, 1855 402

23) 匀点阎甲亚属Carcinops Marseul, 1855 403

(192) 贝氏匀点阎甲Carcinops (Carcinops) penatii Zhang et Zhou, 2007 403

(193) 小匀点阎甲Carcinops (Carcinops) pumilio (Erichson, 1834) 405

(194) 中华匀点阎甲Carcinops (Carcinops) sinensis Lewis, 1909 408

(195) 隐匀点阎甲Carcinops (Carcinops) troglodytes (Paykull, 1811) 409

47. 胸线阎甲属Diplostix Bickhardt, 1921 411

24) 胸线阎甲亚属Diplostix Bickhardt, 1921 412

(196) 卡伦胸线阎甲Diplostix (Diplostix) karenensis (Lewis, 1891) 412

(197) 异胸线阎甲Diplostix (Diplostix) vicaria (Cooman, 1935) 412

48. 厚阎甲属Pachylomalus Schmidt, 1897 415

25) 额线阎甲亚属Canidius Cooman, 1941 415

(198) 缺线厚阎甲Pachylomalus (Canidius) deficiens Cooman, 1933 416

(199) 肌厚阎甲Pachylomalus (Canidius) musculus (Marseul, 1873) 419

49. 隐阎甲属Cryptomalus Mazur, 1993 421

(200) 名豪隐阎甲Cryptomalus mingh Mazur, 2007 421

50. 平阎甲属Platylomalus Cooman, 1948 423

(201) 斯里兰卡平阎甲Platylomalus ceylanicus (Motschulsky, 1863) 424

(202) 折平阎甲Platylomalus inflexus Zhang et Zhou, 2007 426

(203) 门第平阎甲Platylomalus mendicus (Lewis, 1892) 428

(204) 日本平阎甲Platylomalus niponensis (Lewis, 1899) 431

(205) 洋平阎甲Platylomalus oceanitis (Marseul, 1855) 433

(206) 索氏平阎甲Platylomalus sauteri (Bickhardt, 1912) 435

(207) 胸线平阎甲Platylomalus submetallicus (Lewis, 1892) 436

(208) 东京湾平阎甲Platylomalus tonkinensis (Cooman, 1937) 439

(209) 旅平阎甲Platylomalus viaticus (Lewis, 1892) 441

51. 简额阎甲属Eulomalus Cooman, 1937 443

(210) 安普简额阎甲Eulomalus amplipes Cooman, 1937 443

(211) 龙目简额阎甲Eulomalus lombokanus Cooman, 1937 445

(212) 普普简额阎甲Eulomalus pupulus Cooman, 1937 446

(213) 皱简额阎甲Eulomalus rugosus Zhang et Zhou, 2007 448

(214) 赛氏简额阎甲Eulomalus seitzi Cooman, 1941 450

(215) 塔蒂简额阎甲Eulomalus tardipes (Lewis, 1892) 452

(216) 蠕尾简额阎甲Eulomalus vermicipygus Cooman, 1937 454

52. 丽尾阎甲属Paromalus Erichson, 1834 456

26) 丽尾阎甲亚属Paromalus Erichson, 1834 456

(217) 锐角丽尾阎甲Paromalus (Paromalus) acutangulus Zhang et Zhou, 2007 457

(218) 菱丽尾阎甲Paromalus (Paromalus) parallelepipedus (Herbst, 1792) 459

(219) 皮克丽尾阎甲Paromalus (Paromalus) picturatus Kapler, 1999 461

(220) 西藏丽尾阎甲Paromalus (Paromalus) tibetanus Zhang et Zhou, 2007 463

(221) 春丽尾阎甲Paromalus (Paromalus) vernalis Lewis, 1892 465

（八）球阎甲亚科Abraeinae MacLeay, 1819 467

九）球阎甲族Abraeini MacLeay, 1819 468

53. 刺球阎甲属Chaetabraeus Portevin, 1929 468

27) 刺球阎甲亚属Chaetabraeus Portevin, 1929 469

(222) 邦刺球阎甲Chaetabraeus (Chaetabraeus) bonzicus (Marseul, 1873) 469

(223) 合刺球阎甲Chaetabraeus (Chaetabraeus) cohaeres (Lewis, 1898) 471

(224) 粒刺球阎甲Chaetabraeus (Chaetabraeus) granosus (Motschulsky, 1863) 472

(225) 东方刺球阎甲Chaetabraeus (Chaetabraeus) orientalis (Lewis, 1907) 472

28) 马儒阎甲亚属Mazureus Gomy, 1991 474

(226) 等刺球阎甲Chaetabraeus (Mazureus) paria (Marseul, 1856) 474

十）断胸阎甲族Plegaderini Portevin, 1929 475

54. 断胸阎甲属Plegaderus Erichson, 1834 475

29) 断胸阎甲亚属Plegaderus Erichson, 1834 476

(227) 断胸阎甲Plegaderus (Plegaderus) vulneratus (Panzer, 1797) 476

十一）异跗阎甲族Acritini Wenzel, 1944 478

55. 阿拜阎甲属Abaeletes Cooman, 1940 478

(228) 派氏阿拜阎甲Abaeletes perroti (Cooman, 1940) 478

56. 异跗阎甲属Acritus LeConte, 1853 478

30) 密纹阎甲亚属Pycnacritus Casey, 1916 480

(229) 白水隆异跗阎甲Acritus (Pycnacritus) shirozui Hisamatsu, 1965 480

31) 异跗阎甲亚属Acritus LeConte, 1853 481

(230) 库氏异跗阎甲Acritus (Acritus) cooteri Gomy, 1999 481

(231) 驹井异跗阎甲Acritus (Acritus) komai Lewis, 1879 482

(232) 嗜草异跗阎甲Acritus (Acritus) pascuarum Cooman, 1947 484

(233) 梳异跗阎甲Acritus (Acritus) pectinatus Cooman, 1932 485

(234) 瘤胸异跗阎甲Acritus (Acritus) tuberisternus Cooman, 1932 487

十二）条阎甲族Teretriini Bickhardt, 1916 487

57. 条阎甲属Teretrius Erichson, 1834 488

32) 新条阎甲亚属Neoteretrius Kryzhanovskij et Reichardt, 1976 488

(235) 台湾条阎甲Teretrius (Neoteretrius) formosus (Lewis, 1915) 488

(236) 柴田条阎甲Teretrius (Neoteretrius) shibatai ？hara, 2008 489

33) 条阎甲亚属Teretrius Erichson, 1834 489

(237) 太极条阎甲Teretrius (Teretrius) taichii ？hara, 2008 489

（九）腐阎甲亚科Saprininae Blanchard, 1845 489

58. 秃额阎甲属Gnathoncus Jacquelin-Duval, 1858 491

(238) 短胸秃额阎甲Gnathoncus brevisternus Lewis, 1907 492

(239) 迪斯秃额阎甲长缝亚种Gnathoncus disjunctus suturifer Reitter, 1896 492

(240) 克氏秃额阎甲Gnathoncus kiritshenkoi Reichardt, 1930 494

(241) 小齿秃额阎甲Gnathoncus nannetensis (Marseul, 1862) 496

(242) 烟秃额阎甲Gnathoncus nidorum Stockmann, 1957 498

(243) 波氏秃额阎甲Gnathoncus potanini Reitter, 1896 498

(244) 圆秃额阎甲Gnathoncus rotundatus (Kugelann, 1792) 499

(245) 半缘秃额阎甲Gnathoncus semimarginatus Bickhardt, 1920 501

59. 腐阎甲属Saprinus Erichson, 1834 502

34) 狭胸阎甲亚属Phaonius Reichardt, 1941 504

(246) 法老腐阎甲Saprinus (Phaonius) pharao Marseul, 1855 504

35) 腐阎甲亚属Saprinus Erichson, 1834 507

(247) 阿登腐阎甲Saprinus (Saprinus) addendus Dahlgren, 1964 507

(248) 埃腐阎甲Saprinus (Saprinus) aeneolus Marseul, 1870 507

(249) 双斑腐阎甲Saprinus (Saprinus) biguttatus (Steven, 1806) 510

(250) 圆斑腐阎甲Saprinus (Saprinus) bimaculatus Dahlgren, 1964 512

(251) 蓝斑腐阎甲指名亚种Saprinus (Saprinus) caerulescens caerulescens (Hoffmann, 1803) 514

(252) 蓝斑腐阎甲点胸亚种Saprinus (Saprinus) caerulescens punctisternus Lewis, 1900 515

(253) 中央腐阎甲Saprinus (Saprinus) centralis Dahlgren, 1971 518

(254) 金泽腐阎甲Saprinus (Saprinus) chalcites (Illiger, 1807) 518

(255) 齐腐阎甲Saprinus (Saprinus) concinnus (Gebler, 1830) 521

(256) 缓腐阎甲Saprinus (Saprinus) dussaulti Marseul, 1870 523

(257) 丽斑腐阎甲Saprinus (Saprinus) flexuosofasciatus Motschulsky, 1845 524

(258) 额纹腐阎甲Saprinus (Saprinus) frontistrius Marseul, 1855 526

(259) 寒鸦腐阎甲Saprinus (Saprinus) graculus Reichardt, 1930 528

(260) 哈氏腐阎甲Saprinus (Saprinus) havajirii Kapler, 2000 530

(261) 喜马拉雅腐阎甲Saprinus (Saprinus) himalajicus Dahlgren, 1971 531

(262) 污腐阎甲Saprinus (Saprinus) immundus (Gyllenhal, 1827) 531

(263) 铜泽腐阎甲Saprinus (Saprinus) intractabilis Reichardt, 1930 533

(264) 日本腐阎甲Saprinus (Saprinus) niponicus Dahlgren, 1962 535

(265) 丽鞘腐阎甲Saprinus (Saprinus) optabilis Marseul, 1855 538

(266) 扭斑腐阎甲Saprinus (Saprinus) ornatus Erichson, 1834 540

(267) 派腐阎甲Saprinus (Saprinus) pecuinus Marseul, 1855 543

(268) 平盾腐阎甲Saprinus (Saprinus) planiusculus Motschulsky, 1849 545

(269) 侧斑腐阎甲Saprinus (Saprinus) quadriguttatus (Fabricius, 1798) 548

(270) 谢氏腐阎甲Saprinus (Saprinus) sedakovii Motschulsky, 1860 550

(271) 半线腐阎甲Saprinus (Saprinus) semistriatus (Scriba, 1790) 553

(272) 斯博腐阎甲Saprinus (Saprinus) spernax Marseul, 1862 555

(273) 灿腐阎甲Saprinus (Saprinus) splendens (Paykull, 1811) 556

(274) 斯达腐阎甲Saprinus (Saprinus) sternifossa Müller, 1937 558

(275) 淡黑腐阎甲Saprinus (Saprinus) subcoerulus Thérond, 1978 561

(276) 细纹腐阎甲欧亚亚种Saprinus (Saprinus) tenuistrius sparsutus Solskiy, 1876 561

60. 半腐阎甲属Hemisaprinus Kryzhanovskij, 1976 564

(277) 墨半腐阎甲Hemisaprinus subvirescens (Ménétries, 1832) 564

61. 连窝阎甲属Chalcionellus Reichardt, 1932 566

(278) 阿莫连窝阎甲Chalcionellus amoenus (Erichson, 1834) 567

(279) 布氏连窝阎甲指名亚种Chalcionellus blanchii blanchii (Marseul, 1855) 569

(280) 布氏连窝阎甲陶芮亚种Chalcionellus blanchii tauricus (Marseul, 1862) 571

(281) 西伯利亚连窝阎甲Chalcionellus sibiricus Dahlgren, 1969 571

(282) 土耳其连窝阎甲Chalcionellus turcicus (Marseul, 1857) 571

62. 多纹阎甲属Pholioxenus Reichardt, 1932 573

(283) 多纹阎甲Pholioxenus orion Reichardt, 1932 574

63. 连线阎甲属Hypocacculus Bickhardt, 1914 574

36) 连线阎甲亚属Hypocacculus Bickhardt, 1914 575

(284) 光额连线阎甲Hypocacculus (Hypocacculus) spretulus (Erichson, 1834) 575

64. 皱额阎甲属Hypocaccus Thomson, 1867 577

37) 皱额阎甲亚属Hypocaccus Thomson, 1867 578

(285) 巴西皱额阎甲Hypocaccus (Hypocaccus) brasiliensis (Paykull, 1811) 578

(286) 多芮皱额阎甲Hypocaccus (Hypocaccus) dauricus Reichardt, 1930 578

(287) 皱额阎甲Hypocaccus (Hypocaccus) rugifrons (Paykull, 1798) 580

(288) 辛氏皱额阎甲Hypocaccus (Hypocaccus) sinae (Marseul, 1862) 583

38) 滑背阎甲亚属Baeckmanniolus Reichardt, 1926 585

(289) 变线皱额阎甲指名亚种Hypocaccus (Baeckmanniolus) varians varians (Schmidt, 1890) 585

(290) 变线皱额阎甲大陆亚种Hypocaccus (Baeckmanniolus) varians continentalis (Reichardt, 1941) 587

39) 粗额阎甲亚属Nessus Reichardt, 1932 588

(291) 洁皱额阎甲Hypocaccus (Nessus) asticus Lewis, 1911 588

(292) 巴鲁皱额阎甲Hypocaccus (Nessus) balux Reichardt, 1932 590

(293) 沙漠皱额阎甲Hypocaccus (Nessus) eremobius Reichardt, 1932 591

(294) 蒙古皱额阎甲Hypocaccus (Nessus) mongolicus Reichardt, 1932 591

(295) 虎皱额阎甲指名亚种Hypocaccus (Nessus) tigris tigris (Marseul, 1862) 592

65. 稀阎甲属Reichardtiolus Kryzhanovskij, 1959 593

(296) 杜芮稀阎甲Reichardtiolus duriculus (Reitter, 1904) 593

66. 新厚阎甲属Eopachylopus Reichardt, 1926 594

(297) 溪新厚阎甲Eopachylopus ripae (Lewis, 1885) 594

（十）柱阎甲亚科Trypeticinae Bickhardt, 1913 595

67. 柱阎甲属Trypeticus Marseul, 1864 595

(298) 沟额柱阎甲Trypeticus canalifrons Bickhardt, 1913 597

(299) 裂吻柱阎甲Trypeticus fissirostrum Zhang et Zhou, 2007 599

(300) 森柱阎甲Trypeticus nemorivagus Lewis, 1892 601

(301) 索氏柱阎甲Trypeticus sauteri Bickhardt, 1913 602

(302) 猎柱阎甲Trypeticus venator (Lewis, 1884) 603

(303) 云南柱阎甲Trypeticus yunnanensis Zhang et Zhou, 2007 605

参考文献 609

英文摘要 646

中名索引 680

学名索引 687

《中国动物志》已出版书目 703

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